These are for personal use only and not for commercial use or redistribution. An asterisk (*) indicates someone who was a student or post-doc in my lab at the time of the research. You can also view these publications grouped by specific lines of research.


Barclay, P., & Mishra, S. (in press). The psychology of relative state, desperation, and violence: a commentary on de Courson et al. Accepted for publication at Proceedings of the Royal Society of London B.

Arnocky, S., Davis, A.C., Barclay, P., Brittain, H., & Vaillancourt, T. (in press). Actually it does: Errors in “Altruism does not predict mating success in humans: a direct replication.” Forthcoming in Evolutionary Behavioral Sciences.

Maurice, G., Barclay, P., Small, T.A., & Giguère, B. (in press). Do household political campaign signs help win vote share? Forthcoming at Canadian Journal of Political Science.

van Leeuwen, F., Inbar, Y., Bang Petersen, M., Aarøe, L., Barclay, P., Barlow, F.K., de Barra, M., Becker, D.V., Borovoi, L., Choi, J.A., Consedine, N.S., Conway, J.R., Conway, P., Adoric, V.C., Demirci, D.E., Fernández, A.M., Ferreira, D.C.S., Ishii, K., Jakšić, I., Ji, T., Jonaityte, I., Lewis, D.M.G., Li, N.P., McIntyre, J.C., Mukherjee, S., Park, J.H., Pawlowski, B., Pizarro, D., Prokop, P., Prodromitis, G., Rantala, M.J., Reynolds, L.M., Sandin, B., Sevi, B., Srinivasan, N., Tewari, S., Yong, J.C., Žeželj, I., Tybur, J.M. (2023). Disgust sensitivity relates to attitudes toward gay men and lesbian women across 31 nations. Group Processes and Intergroup Relations, 26(3), 629-651. [LINK]

Dhaliwal, N., Martin, J.W., Barclay, P., & Young, L.L. (2022). Signaling benefits of partner choice decisions. Accepted for publication in Journal of Experimental Psychology: General, 151(6), 1466-1472. [LINK] [PDF]

Batistoni, T., Barclay, P., & Raihani, N. (2022). Third-party punishers do not compete to be chosen as partners in an experimental game. Proceedings of the Royal Society of London B289(1966), 20211773. [LINK] [PDF]

*Rotella, A., *Sparks, A.M., Mishra, S., & Barclay, P. (2021). No effect of ‘watching eyes’: An attempted replication and extension investigating individual differences. PLoS ONE, 16(10), e0255531. [LINK] [PDF]

Barclay, P., Bliege Bird, R., Roberts, G., & Számadó, S. (2021). Cooperating to show that you care: costly helping as an honest signal of fitness interdependence. Philosophical Transactions of the Royal Society B, 376, 20200292. [LINK] [PDF]

Roberts, G., Raihani, N., Bshary, R., Manrique, H., Farina, A., Samu, F., & Barclay, P. (2021). The benefits of being seen to help others: indirect reciprocity and reputation-based partner choice. Philosophical Transactions of the Royal Society B, 376, 20200290. [LINK] [PDF]

Manrique, H.M., Zeidler, H., Roberts, G., Barclay, P., Walker, M., Samu, F., Farina, A., Bshary, R., & Raihani, N. (2021). The psychological foundations of reputation-based cooperation. Philosophical Transactions of the Royal Society B, 376, 20200287. [LINK] [PDF]

Wu, J., Számadó, S., Barclay, P., Beersma, B., Dores Cruz, T.D., Lo Iacono, S., Nieper, A.S., Peters, K., Przepiorka, W., Tiokhin, L., & Van Lange, P.A.M. (2021). Honesty and dishonesty in gossip strategies: a fitness interdependence perspective. Philosophical Transactions of the Royal Society B, 376, 20200300. [LINK]

Barclay, P., & Barker, J. (2020). Greener than thou: people who protect the environment are more cooperative, compete to be environmental, and benefit from reputation. Journal of Environmental Psychology, 72, 101441. [LINK] [PDF]

Barclay, P., & Benard, S. (2020). The effects of social versus asocial threats on group cooperation and manipulation of perceived threats. Evolutionary Human Sciences, 2, e54. [LINK] [PDF]

Benard, S., & Barclay, P. (2020). Democratic competition for rank increases both cooperation and deception in small groups. Social Science Quarterly, 101(7), 2413-2436. [LINK] [PDF]

Barclay, P. Barclay, P. (2020). Reciprocity creates a stake in one’s partner, or why you should cooperate even when anonymous. Proceedings of the Royal Society of London B, 287, 20200819. [LINK] [PDF]

*Rotella, A., & Barclay, P. (2020). Failure to replicate moral licensing and moral cleansing in an online experiment. Personality and Individual Differences, 161, 109967. [LINK] [PDF]

*Rotella, A., *Fogg, C., Mishra, S., & Barclay, P. (2019). Measuring delay discounting in a crowdsourced sample: an exploratory study. Scandinavian Journal of Psychology, 60, 520-527. [LINK] [PDF]

Ho, J.L., Powell, D. M., Barclay, P., & Gill, H. (2019). The influence of competition on motivation to fake in employment interviews. Journal of Personnel Psychology, 18(2), 95-105. [LINK] [PDF]

*Larney, A., *Rotella, A, & Barclay, P. (2019). Stake size effects in Ultimatum Game and Dictator Game offers: a meta-analysis. Organizational Behavior and Human Decision Processes, 151, 61-72. [LINK] [PDF]

Barclay, P., *Mishra, S., & *Sparks, AM. (2018). State-dependent risk-taking. Proceedings of the Royal Society of London B, 285, 20180180. [LINK] [PDF]

*O’Connor, J. J. M., & Barclay, P. (2018). High voice pitch mitigates the aversiveness of antisocial cues in men’s speech. British Journal of Psychology, 109, 812-829. [LINK] [PDF]

*Pleasant, A., & Barclay, P. (2018). Why hate the good guy? Antisocial punishment of high cooperators is higher when people compete to be chosen. Psychological Science, 29(6), 868-876. [LINK]

Lightner, A.D., Barclay, P., & Hagen, E.H. (2017). Radical framing effects in the ultimatum game: the impact of explicit culturally transmitted frames on economic decision making. Royal Society Open Science, 4, 170453. [LINK] [PDF]

*O’Connor, J., & Barclay, P. (2017). The influence of voice pitch on perceptions of trustworthiness across social contexts. Evolution and Human Behavior, 38(4), 506-512. [LINK] [PDF]

Montano, K. J., Tigue, C. C., Isenstein, S. G. E., Barclay, P., & Feinberg, D. (2017). Men’s voice pitch influences women’s trusting behavior. Evolution and Human Behavior, 38(3), 293-297. [LINK] [PDF]

Driscoll, R. L., Barclay, P., & Fenske, M. (2017). To be spurned no more: The affective and behavioural consequences of social and non-social rejection. Psychonomic Bulletin and Review, 24, 566-573. [LINK] [PDF]

Arnocky, S., Piché, T., Albert, G., Ouellette, D., & Barclay, P. (2017) Altruism predicts mating success in humans. British Journal of Psychology, 108, 416-435. [LINK] [PDF]

*Mishra, S., Barclay, P., & *Sparks. (2017). The relative state model: Integrating need-based and ability-based pathways to risk-taking. Personality and Social Psychological Review, 21(2), 176-198. [LINK] [PDF]

Barclay, P. (2017). Bidding to commit: an experimental test of the benefits of commitment under moderate degrees of conflict. Evolutionary Psychology, 15(1), 1-8. [LINK] [PDF]

Tybur, J. M., Inbar, Y., Aarøe, L., Barclay, P., Barlow, F.K., de Barra, M., Becker, D.V., Borovoi, L., Choi, I., Choi, J.A., Consedine, N.S., Conway, A., Conway, J.R., Conway, P., Cubela Adoric, V., Demirci, E., Fernández, A.M., Ferreira, D.C.S., Ishii, K., Jakšić, I., Ji, T., van Leeuwen, L., Lewis, D.M.G., Li, N.P., McIntyre, J.C., Mukherjee, S., Park, J., Pawlowski, B., Petersen, M.B., Pizarro, D., Prodromitis, G., Prokop, P., Rantala, M.J., Reynolds, L.M., Sandin, B., Sevi, B., de Smet, D., Srinivasan, N., Tewari, S., *Wilson, C., Young, J.C., & Žeželj, I. (2016). Parasite stress and pathogen avoidance relate to distinct dimensions of political ideology across 30 nations. Proceedings of the National Academy of Sciences of the USA, 113(44), 12408-12413. [LINK] [PDF]

Raihani, N., & Barclay, P. (2016). Exploring the trade-off between quality and fairness in human partner choice. Royal Society Open Science, 3, 160510. [LINK] [PDF]

Barker, J., & Barclay, P. (2016). Local competition increases people’s willingness to harm others, including group members. Evolution and Human Behavior37, 315-322. [LINK] [PDF]

Barclay, P., & Raihani, N. (2016). Partner choice versus punishment in human prisoner’s dilemmas. Evolution and Human Behavior37, 263-271. [LINK] [PDF]

*Sparks, A., Burleigh, T., & Barclay, P. (2016). We can see inside: Accurate predictions of Prisoner’s Dilemma decisions after a brief face-to-face interaction. Evolution and Human Behavior37, 210-216. [LINK] [PDF]

Barclay, P. (2016). Biological markets and the effects of partner choice on cooperation and friendship. Current Opinion in Psychology, 7, 33-38. [LINK] [PDF]

*Sparks, A., & Barclay, P. (2015). No effect on condemnation of short or long exposure to eye images. Letters on Evolutionary Behavioral Science, 6(2), 13-16. [LINK] [PDF]

Barclay, P., & *Stoller, B. (2014). Local competition sparks concern for fairness in the ultimatum game. Biology Letters, 10, 20140213. [LINK] [PDF]

*Mishra, S., Barclay, P., & Lalumière, M.L. (2014). Competitive disadvantage facilitates risk taking. Evolution and Human Behavior, 35, 126-132[LINK] [PDF]

Barclay, P., & Benard, S. (2013). Who cries wolf, and when: manipulation of perceived threats to preserve rank in cooperative groups. PLOS ONE, 8(9), e73863. doi:10.1371/journal.pone.0073863. [LINK with PDF]

Barclay, P. (2013). Strategies for cooperation in biological markets, especially for humans. Evolution & Human Behavior, 34(3), 164-175. [LINK] [PDF] * This paper was one of the most cited articles in Evolution & Human Behavior from 2011-2016.

*Sparks, A., & Barclay, P. (2013). Eyes increase generosity, but not for long: the limited effect of a false cue. Evolution & Human Behavior, 34, 317-322. [LINK] [PDF] * This paper was also one of the most cited articles in Evolution & Human Behavior since 2011.

Barker, J., Barclay, P., & Reeve, H.K. (2013). Competition over personal resources favors contributions to shared resources in human groups. PLOS ONE, 8(3), e58826. [LINK with PDF]

Barclay, P. (2012). Harnessing the power of reputation: strengths and limits for promoting cooperative behaviours. Evolutionary Psychology, 10(5), 868-883. [LINK] [PDF]

Barclay, P., & Reeve, H.K. (2012). The varying relationship between helping and individual quality. Behavioral Ecology, 23(4), 693-698. [LINK] [PDF]

Barker, J., Barclay, P., & Reeve, H.K. (2012). Within-group competition reduces cooperation and payoffs in human groups. Behavioral Ecology, 23(4), 735-741. [LINK] [PDF]

Barclay, P. (2011). Competitive helping increases with the size of biological markets and invades defection. Journal of Theoretical Biology, 281, 47-55. [LINK] [PDF]

Barclay, P. (2010). Altruism as a courtship display: Some effects of third-party generosity on audience perceptions. British Journal of Psychology, 101, 123-135. [LINK] [PDF]

Kiyonari, T., & Barclay, P. (2008). Cooperation in social dilemmas: free-riding may be thwarted by second-order rewards rather than punishment. Journal of Personality and Social Psychology, 95(4), 826-842. [LINK] [PDF]

Krupp, D. B., DeBruine, L. M., & Barclay, P. (2008). A cue of kinship promotes cooperation for the public good. Evolution & Human Behavior, 29, 49-55. [LINK] [PDF]

Barclay, P. (2008). Enhanced recognition of defectors depends on their rarity. Cognition, 107, 817-828. [LINK] [PDF]

Barclay, P., & Willer, R. (2007). Partner choice creates competitive altruism in humans. Proceedings of the Royal Society of London Series B, 274, 749-753. [LINK] [PDF]

Barclay, P. (2006). Reputational benefits for altruistic punishment. Evolution and Human Behavior, 27, 325-344. [LINK] [PDF]

Barclay, P., & Lalumière, M. (2006). Do people differentially remember cheaters? Human Nature, 17(1), 98-113. [LINK] [PDF]

Barclay, P. (2004). Trustworthiness and Competitive Altruism Can Also Solve the “Tragedy of the Commons”. Evolution & Human Behavior, 25(4), 209-220. [LINK] [PDF]


*Kafashan, S., *Sparks, A., *Rotella, A., & Barclay, P. (2016). Why Heroism Exists: Evolutionary Perspectives on Extreme Helping. In S.T.Allison, G.R.Goethals, & R.M.Kramer (Eds.) The Handbook of Heroism and Heroic Leadership, pp. 36-57. Routledge. [PDF]

Barclay, P. (2015). Reputation. In D. Buss (Ed.) Handbook of Evolutionary Psychology (2nd Ed.), pp. 818-828. Hoboken, NJ: J. Wiley & Sons. [PDF]

Barclay, P., & Van Vugt, M. (2015). The evolutionary psychology of human prosociality: adaptations, mistakes, and byproducts. In D. Schroeder & W. Graziano (Eds.) Oxford Handbook of Prosocial Behavior, pp. 37-60. Oxford, UK: Oxford University Press. [PDF] * Note: most of this chapter was later reused with permission in the authored book “Social Dilemmas: The Psychology of Human Cooperation” by P. Van Lange, D. Balliet, C. D. Parks, & M. Van Vugt, published in 2014 by Oxford University Press. I am first author of the corresponding chapter there.

*Kafashan, S., *Sparks, A., Griskevicius, V., & Barclay, P. (2014). Prosocial behaviour and social status. In J. T. Cheng, J. L. Tracy, & C. Anderson (Eds.) The Psychology of Social Status, pp. 139-158. New York, NY: Springer. [PDF]

Barclay, P., & Kiyonari, T. (2014). Why sanction? Functional causes of punishment and reward. In P. Van Lange, B. Rockenbach, & T. Yamagishi (Eds.) Social Dilemmas: New Perspectives on Reward and Punishment, pp. 182-196. Oxford, UK: Oxford University Press. [PDF]

Barclay, P. (2011). The evolution of charitable behaviour and the power of reputation. In C. Roberts (Ed.) Applied Evolutionary Psychology, pp. 149-172. Oxford, UK: Oxford University Press. [PDF]

Barclay, P. (2010). Reputation and the Evolution of Generous Behavior. Nova Science Publishers, Hauppauge, NY. * Note: this is essentially a very lengthy book chapter based on my PhD thesis introduction & discussion, but the publisher released it as a stand-alone book. [PDF]


Barclay, P., *Yerman, S., & *Twardus, O. (2023). Fearful apes or emotional cooperative breeders? Behavioral and Brain Sciences, 46, e53.

*Rotella, A., Sparks, A.M., & Barclay, P. (2020). Feelings of obligation are valuations of signaling-mediated social payoffs. Behavioral and Brain Sciences, 43, e85.

Karabegović, M., *Rotella, A., & Barclay, P. (2018). Broadening the role of “self-interest” in Folk-Economic Beliefs. Behavioral and Brain Sciences, 41, e174.

Barclay, P. (2017). Attractiveness biases are the tip of the iceberg in biological markets. Behavioral and Brain Sciences, 40, 17-18.

Barclay, P., & Krupp, D. B. (2016). The burden of proof for a cultural group selection account. Behavioral and Brain Sciences39, 21-22. [PDF]

Krupp, D. B., Kim, J., Taylor, P., & Barclay, P. (2014). Cooperation and competition in large classrooms. Toronto: Higher Education Quality Council of Ontario. [PDF]

*Sparks, A., *Mishra, S., & Barclay, P. (2013). Fundamental freedoms and the psychology of threat, bargaining, and inequality. Behavioral and Brain Sciences, 36, 36-37. [PDF]

Barclay, P. (2013). Pathways to Abnormal Revenge and Forgiveness. Behavioral and Brain Sciences, 36(1), 17-18. [PDF]

Barclay, P. (2012). Proximate and ultimate causes of Strong Reciprocity and punishment. Behavioral and Brain Sciences, 35(1), 16-17. [PDF]

Krupp, D. B., & Barclay, P. (2010). Margo Wilson (1942-2009). Journal of Evolutionary Psychology, 8(1), 1-3. [PDF]

Barclay, P. (2008). Using the hatchet and burying it afterwards – A review of “Beyond revenge: The evolution of the forgiveness instinct”. Invited book review for Evolution & Human Behavior, 29(6), 450-451. [PDF]

Barclay, P. (2006). Dissertation abstract: Reputational benefits of altruism and altruistic punishment. Experimental Economics, 9(2), 181-182.

Krupp, D.B., Barclay, P., Daly, M., Kiyonari, T., Dingle, G., & Wilson, M. (2005). Let’s add some psychology (and maybe even some evolution) to the mix. Behavioral & Brain Sciences, 28(6), 828-829. [PDF]

Barclay, P., & Daly, M. (2003). Humans should be individualistic and utility-maximizing, but not necessarily “rational”. Behavioral & Brain Sciences, 26(2), 154-155. [PDF]